What do Macaroni penguins eat besides fish?

Macaroni penguins primarily eat fish. They are not known to eat other things besides fish.

Here's why:

* Specialized Diet: Macaroni penguins have evolved to be efficient fish eaters. Their beaks and tongues are perfectly adapted for catching and swallowing fish.

* Ocean Environment: Macaroni penguins live in the Southern Ocean, a rich environment with abundant fish populations.

* Limited Options: Unlike some other penguin species, Macaroni penguins don't have access to other food sources like krill or squid in large quantities.

While they might occasionally ingest a small invertebrate along with their fish, their diet is overwhelmingly based on fish.